Description:
This grass is a summer annual that becomes 4-16' tall at maturity.
The culm is stout, terete, light green, glabrous, and unbranched.
The interior of the culm is solid, containing a white pith. Alternate
leaves occur along the entire length of the culm, their blades becoming
largest toward the lower-middle or middle of the culm. The leaf blades
are 10-40" long and ¾–4½" across; they are ascending toward their
bases,
but arching toward their tips, and rather floppy overall. The leaf
blades are linear to linear-lanceolate in shape, while their margins
are entire (without conspicuous teeth) and often ciliate near the bases
of the leaf blades. In addition, the leaf blades may have narrow
longitudinal wrinkles, and they are sometimes vertically wavy (up and
down). The leaf blade surfaces are medium green, hairless or nearly so,
and rather
shiny; they have prominent central veins that are whitened, especially
toward their bases. The leaf bases strongly clasp the culm, often
forming rounded auricles (ear-like basal lobes). The open leaf sheaths
are light green to light reddish green, relatively tight, and mostly
glabrous, except along their margins, where they are hairy and narrowly
membranous. The ligules are membranous, while the ring-like nodes are
slightly swollen and glabrous.
This grass produces a single terminal
staminate inflorescence (called the 'tassel') and several axillary
pistillate inflorescences (called the 'ears'). The staminate
inflorescence is typically about 1' long and a little less across,
consisting of a panicle of spike-like racemes; it is greenish yellow
during the blooming period. This panicle has a straight central rachis
and several lateral racemes originating from this rachis that
are ascending to arching. The central axis is covered with about 4
rows
of staminate spikelets along most of its length, while the lateral
racemes are covered with about 2 rows of staminate spikelets. The
staminate spikelets are about 9-14 mm. in length, ellipsoid-lanceoloid
in shape, and slightly flattened. Each staminate spikelet consists
of a pair of outer glumes, a pair of membranous lemmas, and a pair of
staminate florets that are separated by membranous paleae.
Each staminate floret has 3 anthers that are yellow, elongated in
shape, and relatively large. During the period of bloom, the axillary
pistillate inflorescences are 6-12" long, 1-3½" across,
cylindrical-lanceoloid in shape, and semi-erect; they occur on short
stout peduncles from the axils of the lower-middle to upper leaves,
where they are solitary. Each pistillate inflorescence is mostly
covered by modified sheaths (called the 'husks') that are light green,
glabrous (or nearly so), and longitudinally wrinkled; some of these
sheaths may have diminutive blades that are medium green. The central
rachis of each pistillate inflorescence (called the 'cob') is thick,
terete, and slightly woody; surrounding this rachis are 8-32 rows of
pistillate spikelets that are densely bunched together.
Each pistillate
spikelet has one fertile floret and one infertile (aborted) floret; the
pistillate spikelet has a pair of short fleshy glumes, a short
membranous lemma, a short membranous palea, and an ovary with a pair of
styles. The two styles are fused together, except near their tips; they
are long, silky, and filamentous. The filamentous styles are exerted
from the tip of
the pistillate inflorescence and stigmatic along their sides.
Collectively, these styles are referred
to as the 'silk' of the ear (pistillate inflorescence). The blooming
period occurs from mid- to late summer, lasting about 1-2 weeks for a
colony of plants. The florets are cross-pollinated by the wind. While
some limited self-pollination can occur, mature grains (called
'kernels') are more abundant when cross-pollination from neighboring
plants occur. After the blooming period, the pistillate inflorescences
(ears of corn) enlarge in size slightly and the pistillate florets are
replaced by kernels (seeds) that mature during the autumn. Mature
kernels are
either ovoid in shape, or they are obovoid and somewhat flattened in
shape; they are exerted beyond the glumes and hard-coated. The kernels
on an ear of corn can be yellow, white, red, purplish-blue, or
some combination of these colors (as occurs in 'Indian Corn'), although
they are mostly typically yellow. Both ears of corn and their kernels
are
resistant to disarticulation. The root system is coarsely fibrous.
Support roots often develop from the lower nodes of the culm. This
grass reproduces by reseeding itself.
Cultivation: Cultivated
corn usually prefers full sun, mesic levels of moisture, and fertile
loamy soil. Because of its C4 metabolism, cultivated corn grows best in
warm to slightly hot summer weather. Flooded conditions, severe
drought, and infertile soil are detrimental to the growth and
development of this grass. Because its pollen is rather heavy, more
mature kernels on ears of corn are produced when corn plants are
adjacent to each other in relatively large blocks. This arrangement
also reduces the likelihood that individual plants will topple over in
high winds. The size of individual plants can be highly variable. There
are many different kinds of cultivated corn that can vary somewhat in
their growing requirements.
Range
& Habitat: Cultivated corn has probably
naturalized, at one time or
another, in every county of Illinois, although it never persists
outside of cultivation. Current collections of naturalized plants
underestimate the ubiquitousness of this species (see
Distribution
Map).
The ancestor of modern cultivated corn probably originated from
southern Mexico. From there, older strains of cultivated corn spread
northward to include the southern half of North America (including
Illinois) prior to the period of European settlement. Current habitats
of modern naturalized plants include cropland, fallow fields, vacant
lots,
roadsides, areas along railroads, areas around grain elevators, and
areas around bird feeders. Cultivated corn is a major agricultural
crop in Illinois, and it is occasionally cultivated as a source of food
for wildlife in natural areas. This grass prefers sunny habitats with a
history of disturbance where the topsoil has been exposed.
Faunal
Associations: Because cultivated corn is a major
agricultural crop,
there is considerable knowledge about the insects that feed on it.
Examples of such species include the Corn Flea Beetle (
Chaetocnema
pulicaria), Northern Corn Rootworm (
Diabrotica barberi),
Southern Corn
Leaf Beetle (
Myochrous
denticollis), LeConte's Seed Corn Beetle
(
Stenolophus lecontei),
Southern Corn Billbug (
Sphenophorus
callosus),
Maize Billbug (
Sphenophorus
maidis), Corn Blotch Leafminer (
Agromyza
parvicornis), Seed Corn Maggot (
Delia platura),
Common Chinch Bug
(
Blissus leucopterus
leucopterus), Red-shouldered Stink Bug (
Thyanta
custator accerra), Corn Leaf Aphid (
Rhopalosiphum maidis),
Corn Earworm
Moth (
Helicoverpa zea),
European Corn Borer Moth (
Ostrinia
nubilalis), Eufala Skipper (
Lerodea eufala),
Two-striped Grasshopper (
Melanoplus
bivittatus), Migratory Grasshopper
(
Melanoplus sanguinipes),
and Eastern Grass Thrips (
Anaphothrips
obscura). The
Insect Table and
Moth
Table
display more complete
lists of these species.
Cultivated corn is also an important source of food for
many kinds of vertebrate wildlife. The kernels are eaten by many
species
of ducks, upland gamebirds, songbirds, and other birds. This includes
such species as the Sandhill Crane, Canada Goose, Mallard, Northern
Pintail, Greater Prairie Chicken, Ring-necked Pheasant, Redwing
Blackbird, American Crow, Common Grackle, House Sparrow, Lincoln
Sparrow, and Red-headed Woodpecker. The seeds were also eaten by two
extinct birds, the Passenger Pigeon and Carolina Parakeet. The
Bird
Table has a more complete list of these species. Some mammals
also rely
on cultivated corn as a source of food. The kernels are eaten
by the Raccoon, Opossum, Fox Squirrel, Gray Squirrel, House Mouse, and
Prairie Deer Mouse, while the foliage of either seedlings or more more
mature plants is eaten by the White-tailed Deer, Groundhog, and
domesticated cattle (Martin et al., 1951/1961; Whitaker, 1966). There
are also records of the Painted Turtle (
Chrysemys picta)
and Spiny
Softshell (
Trionyx
spiniferus) using corn as a minor food source (Ernst
et al., 1994).
Photographic
Location: A corn field at Meadowbrook Park in Urbana,
Illinois.
Comments:
Cultivated corn (
Zea mays)
has a very distinctive appearance, making it
easy to identify. There are other corn species (
Zea spp.) that are
found in southern Mexico and other areas of Central America, but they
are not normally found in Illinois as either wild or cultivated plants.
These other corn species have much smaller corn ears than cultivated
corn, and the glumes of their corn ears usually enclose the kernels
(pod corn). The closest relative to cultivated corn in Illinois is the
native Eastern Gama Grass (
Tripsacum
dactyloides). This latter grass
produces large leaves that have some resemblance to those of cultivated
corn, and there are some structural similarities in their
inflorescences. However, Eastern Gama Grass differs by producing both
male (staminate) and female (pistillate) flowers on the same terminal
inflorescence, while cultivated corn produces a terminal male
inflorescence (the tassel) and axillary female inflorescences (corn
ears) that are separated from each other. There are several kinds of
cultivated corn that are used to produce field corn, sweet corn,
popcorn, corn flour, and ornamental corn. Cultivated corn has been used
to make such products as corn chips, tortillas, tostados, corn meal,
breakfast cereal, corn syrup, corn oil, distilled liquor, animal feed,
bird seed, and ethyl alcohol in gasoline. Another common name of
cultivated corn is maize.