Description:
This sedge is a summer annual (in Illinois) that becomes 6–24" (15–60
cm.) tall. The culm is light to medium green, bluntly 3-angled, and
hairless.
There are usually 2–3 leaves along the lower one-third of the culm that
have ascending, arching, or widely spreading blades. The leaf blades
are 3–12" (7.5–30 cm.) long and 3–8 mm. across; they are longitudinally
furrowed and often shallowly M-shaped in cross-section. The upper blade
surface is light to medium green and hairless, while the lower blade
surface is
light grayish green or whitish green and often slightly glaucous. The
leaf sheaths are light to medium green and hairless, often turning
brown toward
the base of the plant. The culm terminates in a rather congested umbel
or compound umbel of floral spikes spanning up to 5" across and 4"
tall; there is a whorl of several leafy bracts underneath. There are
typically 1 or 2 sessile spikes and either solitary or clustered spikes
at the tips of 3–6 peduncles (floral stalks) up to 4" long; there are
typically 2–4 spikes per cluster. The peduncles are are ascending to
erect, straight, light to medium green, and hairless.
Individual spikes are ½–1½"
(12.5–37.5 mm.) long with a short-cylindrical shape, resembling
bottle brushes from the linear spikelets pointing in all directions.
The
color of the spikes varies primarily with the maturity of the scales of
the
spikelets; they are initially yellowish or whitish green, becoming
yellowish or whitish brown, and finally reddish brown or deep
brown at maturity. Individual spikelets
are 7.5–12.5 mm. long, 1 mm. across, linear in shape, and
terete
but slightly flattened (subterete). The rachilla (central stalklet) of
each spikelet is jointed and slightly zigzag. Each spikelet has 8–18
scales with perfect florets; the scales are slightly to moderately
overlapping. Individual scales are about 2–2.5 mm. long, oblong-ovate,
and convex along their green to brown central veins; their tips are
somewhat acute and often mucronulate (tapering to very short narrow
points). This latter characteristic can cause the lateral sides of the
spikelets to appear slightly jagged. The lateral sides of the scales
also have 1–3 fine longitudinal ribs (may require 10x magnification to
see). The florets have 3 stamens and a tripartite style. The blooming
period can occur from mid-summer to early autumn, lasting about 2–4
weeks for a colony of plants. The florets are cross-pollinated by the
wind. Afterwards, they are replaced by achenes that 1–1.5 mm. long,
oblongoid-ovoid in shape, and light brown to dark brown. The root
system consists of fibrous roots (in Illinois). In more tropical areas,
this sedge can be a short-lived perennial that occasionally produces
short rhizomes.
Cultivation:
The
preference is partial to full
sun, wet to moist conditions, and soil that is sandy, silty, or mucky.
Most growth and development occurs during the summer. Individual plants
can vary significantly in size depending on when the seeds germinate
and growing conditions.
Range
&
Habitat: Rusty Flatsedge
(Cyperus odoratus) is occasional to locally common throughout Illinois
(see
Distribution
Map). It occurs in many areas of North America,
South America, Africa, Australia, south Asia, and the Middle East. In
Illinois, habitats include marshes, shorelines along ponds and rivers,
openings in floodplain woodlands, sandbars, mudflats, margins of
sloughs, prairie swales, damp areas along fields, roadside ditches, and
meridians of highways. Areas with a history of disturbance are
preferred. Rusty Flatsedge has some tolerance to the higher levels of
salt along roadsides.
Faunal
Associations: Insects that feed on
Rusty Flatsedge (Cyperus odoratus) and other flatsedges (Cyperus spp.)
include the larvae of
Bactra
verutana (Javelin Moth) and
Spodoptera
frugiperda (Fall Armyworm), larvae of
Euphyes vestris (Dun
Skipper),
Ischnodemus rufipes (a
chinch bug),
Sphenophorus aequalis (Clay-colored
Billbug) and other
Sphenophorus
spp. (billbugs), larvae of
Plantella caudata (Sedge
Gall Midge),
Sipha
flava (Yellow Sugar Cane Aphid) and other aphids, and
Pseudococcus
sorghiellus (Trochanter Mealybug) and other mealybugs
(Frick &
Wilson, 1978; Barlow & Kuhar, 2009; Wagner, 2005; Hoffman,
1996;
Majka et al., 2007; Vaurie, 1983; Felt, 1917; Blackman &
Eastop,
2013; ScaleNet website, 2014). The seeds of Rusty Flatsedge are an
important food source for some ducks, including the Northern Pintail,
American Wigeon, Green-winged Teal, and Mallard. Another waterfowl, the
Canada Goose, feeds on the foliage (Havera, 1999). Other birds that eat
the
seeds of flatsedges include the American Tree Sparrow, Bobolink, Sora,
Virginia Rail, and Wilson's Snipe (Martin et al., 1951/1961). Among
mammals, Muskrats eat the culms of flatsedges to a minor extent
(Hamerstrom & Blake, 1939). Their foliage is considered
low-value
browse to White-tailed Deer; it is also eaten by goats and cattle
occasionally (Ramachandran & Soosairaj, 2014).
Photographic
Location: A muddy shoreline along Crystal Lake in Urbana,
Illinois. The
photographed plant is the typical variety,
Cyperus odoratus odoratus.
Comments:
Another scientific name of this flatsedge is
Cyperus ferruginescens,
and another common name is Fragrant Flatsedge. Rusty Flatsedge (Cyperus
odoratus) is
another fairly common bottle brush flatsedge that prefers disturbed
areas. It is distinct from many other flatsedges (Cyperus spp.) in
Illinois by the tendency of the rachillas of its spikelets to break
apart into segments to which the floral scales and achenes have been
attached.
Normally, the floral scales and achenes fall from their spikelets
without the rachillas breaking apart into segments. Rusty Flatsedge has
longer scales and achenes than Red-rooted Flatsedge (Cyperus
erythrorhizos), but they are shorter than the scales and achenes of
Straw-colored Flatsedge (Cyperus strigosus). Another difference is that
the spikelets of Rusty Flatsedge are more terete in cross-section than
the spikelets of other flatsedges, which are usually more flattened.
Across its worldwide distribution, Rusty Flatsedge is rather variable
in its appearance. In particular, the extent to which its scales
overlap each other along the spikelets can vary. For the typical
variety of this plant,
Cyperus
odoratus odoratus, the scales moderately
overlap each other, while for
Cyperus
odoratus englemannii, the scales
barely overlap each other. Both of these varieties can be found in
Illinois; some authors prefer to categorize them as distinct species.