Description:
This sod-forming perennial grass is 2-6' high. The unbranched culms are
more or less erect, terete, and glabrous; they are largely covered
by the sheaths of the leaves. Along the entire length of each mature
culm,
there are 5-10 alternate leaves. The rather floppy leaf blades along
the culm are ascending to widely spreading and often arched. Individual
blades are 4-10 mm. across and 8-24" long; they are grayish green
or grayish blue, linear in shape, entire along their margins, and
hairless. The tips of the blades are long and slender, their margins
often rolling into narrow points. The bases of the blades near the culm
are often yellowish. The open leaf sheaths are grayish green or grayish
blue and they are glabrous or pubescent; usually there are some white
hairs toward the apex of each sheath, while the lower sheath is
typically hairless. The ligules have rings of short white hairs. A
fertile culm terminates in a panicle of spikelets about 6-18" long. In
the typical variety of Sand Reed, the panicle has a spike-like
appearance from lateral branches that are erect and appressed; in
var.
magna of Sand Reed, the panicle is more open and
obpyramidal to rhomboid
in outline from primary lateral branches that are ascending.
Secondary lateral
branches are usually appressed along the lengths of primary lateral
branches. The branches and central rachis of the panicle are straight,
very slender, and glabrous, although the tip of the rachis sometimes
nods. One-flowered spikelets develop from short slender pedicels. Each
narrow spikelet is 6-7.5 mm. long and greenish white, consisting of a
pair of glumes, a single lemma, a single membranous palet, and a
perfect floret with 3 anthers and an ovary with a pair of stigmata. The
glumes,
lemma, and palet are narrowly lanceolate in shape and glabrous, except
for a small tuft of fine hairs at the base of the lemma and palet. The
glumes and lemma are single-veined, and their tips are acute
(tapering to a point), but not awned. The glumes are
unequal in length (3-5 mm. long versus 5-7.5 mm. long), while the lemma
is typically 4.5-7.0 mm. long. The blooming period occurs from
mid-summer to early
fall. The florets are cross-pollinated by the wind. Afterwards, the
florets are
replaced by slender pale grains about 2.5-4.5 mm. long, which become
mature during the fall. Disarticulation of the spikelets is above the
glumes. The root system is fibrous and long-rhizomatous, often
extending to a depth of 4-10'. Occasionally, clonal colonies
of plants are produced.
Cultivation:
The preference is full or
partial sun, mesic to dry conditions, and sandy soil (from coarse sand
to sandy loam). Most growth and development occur during the summer.
Tolerance of drought is excellent.
Range
& Habitat: The
native Sand Reed is occasional in sandy areas in the northern half of
Illinois, while in the rest of the state it is rare or absent (see
Distribution
Map).
Habitats include upland sand prairies, sandy hill
prairies, sand dunes, upper areas of beaches along Lake
Michigan, upland sandy savannas,
and openings in sandy woodlands. Sand Reed is
one of the grasses that stabilizes sand dunes and sand
blowouts. In
upland wooded areas, it is associated with oak trees (especially Black
Oak). In some upland sand prairies, this is the dominant grass.
Faunal Associations: Sand Reed is the host plant of a
leafhopper,
Flexamia
grammica, and the adults of a spittlebug,
Philaenarcys killa.
Some species of grasshopper that are often found
in barren sandy areas also feed on Sand Reed; they include such
species as
Eritettix
simplex (Velvet-striped Grasshopper),
Melanoplus foedus
(Striped Sand Grasshopper),
Mermiria
bivittata
(Two-striped Mermiria Grasshopper),
Orphulella speciosa
(Pasture Grasshopper), and
Spharagemon
collare (Mottled Sand
Grasshopper); see the
Grasshopper Table
for a more complete list of these species. For hoofed mammalian
herbivores (cattle, horses,
sheep, deer, etc.), Sand Reed provides below-average forage. However,
young plants are more palatable to such herbivores, and this grass will
be grazed, regardless of its stage of development, when more palatable
grasses and broad-leaved plants are hard to find. The Plains Pocket
Gopher eats the leaves, stems (culms), and rhizomes. This burrowing
animal is found in open sandy areas along the Illinois River and in the
Kankakee sand region of Illinois, where Sand Reed occurs.
Photographic
Location: An upland sandy savanna near Lake Michigan at
the Indiana
Dunes State Park in NW Indiana.
Comments:
Sand Reed superficially resembles
Panicum
virgatum
(Switch Grass),
Calamagrostis canadensis
(Bluejoint Grass), and other tall-growing
grasses with single-seeded florets. It differs from most of
these species by the tuft of fine hairs at the bottom of each lemma and
palea, which may require a 10x hand lens to see (after the glumes have
been removed). While Bluejoint Grass shares this characteristic, it
differs from Sand Reed by the presence of awns on its lemmas. Other
distinctive characteristics of Sand Reed are the pale color of its
spikelets (greenish white or light tan), the relatively long length of
its spikelets (6.0-7.5 mm.) among those spikelets that are
single-seeded, and the
presence of only a single prominent vein on each lemma and glume.