Drummond's Goldenrod
Solidago drummondii
Aster family (Asteraceae)
Description:
This herbaceous perennial wildflower overwinters as a rosette of
semi-evergreen leaves. Beginning in mid- to late spring, it bolts to
become about 1¼–3' tall by autumn. The central stem is light green
(sometimes with streaks of light purple), terete, evenly
short-pubescent, erect to ascending, and mostly unbranched,
although short secondary stems often develop from the upper leaf
axils. Alternate leaves occur along the entire length of the central
stem, becoming gradually smaller as they ascend. Individual leaves are
1¼–3¼" long and 1–2¾" across (about 1.2–2 times as long as wide); they
are ovate to broadly ovate with rather flattened serrated teeth along
their
margins. The tips of the leaves are broadly acute, while their bases
taper somewhat abruptly to short winged petioles. The winged margins of
the petioles are widest where they join the leaf blades, but disappear
where they join the stems. The upper leaf surface is medium to dark
green and sparsely short-hairy to nearly glabrous, while the lower leaf
surface is light green and sparsely to moderate short-pubescent. The
lower leaf surface is more pubescent along the undersides of major
veins than between the veins. Leaf venation is pinnate with a prominent
central vein; 2 prominent lateral veins are present on some leaves but
not others.
The central stem terminates in a pyramidal panicle of
flowerheads that is up to 1¼' long and 1' across; the panicle is
usually widest toward the lower bottom. In response to the weight of
this inflorescence, there is a tendency for the entire plant to arch to
one side. Relative to the orientation of the central branch of the
panicle, the lateral branches are mostly ascending or
ascending-arching. The branchlets terminating in flowerheads are up to
1/3" (8 mm.) long and they bend upward to hold the flowerheads in an
erect position, regardless of the orientation of the panicle. Both the
branches and branchlets of the inflorescence are light to medium green,
terete, and densely short-pubescent. Interspersed along the branches of
the inflorescence are
leafy bracts up to 1" long and ¾" across; the leafy bracts are oval,
broadly elliptic, obovate, or ovate in shape, while their margins are
toothless. Along the terminal branchlets, there are a few leafy
bractlets up to
1/8" (3 mm.) long that are green, broadly oblong in shape, and
scale-like in appearance. Both the leafy bracts and leafy bractlets are
medium to dark green and densely short-pubescent along their lower
sides. The flowerheads (including both their involucres and florets)
are about ¼" (5-6 mm.) tall and ¼" (5-6 mm.) across. Each flowerhead
has 4-7 disk florets that are surrounded by 4-6 ray florets.
The corollas of the disk florets are yellow and short-tubular in shape
with 5 arching lobes at their apices. The petaloid rays of the ray
florets are yellow, widely spreading and short-oblong in shape with 1
or 2 shallow notches at their tips. The base of each flowerhead is
surrounded by scale-like phyllaries (floral bracts) in about 3
irregular series. These phyllaries are unequal in size, light green,
oblong with rounded or bluntly acute tips, and appressed to together;
their margins are short-ciliate. Both the disk florets and ray florets
are perfect.
The blooming period occurs during early to
mid-autumn, lasting about one month. Much smaller and sparsely branched
panicles of flowerheads may bloom from short leafy stems that develop
from the axils of the upper leaves. Fertile florets are replaced by
small achenes with sessile tufts of white hair; they are
distributed by the wind. The achenes are about 1.5–2 mm. long and
narrowly bullet-shaped. The root system consists of a short branching
rootstock, sometimes forming a small caudex on older robust plants.
Sometimes clonal plants are produced from short stout rhizomes.
Cultivation:
The preference is full sun to moderate shade, moist to dry conditions,
and calcareous soil containing clay, loam, or rocky material. This
uncommon goldenrod, notwithstanding its restricted range, adapts
readily to flower gardens in a variety of situations.
Range
& Habitat: The native Drummond's
Goldenrod is restricted to the
border counties of southwest Illinois, where it is uncommon (see Distribution
Map).
This goldenrod is a Midwestern endemic plant that is
found primarily in Missouri and Arkansas. Habitats include borders of
wooded bluffs, openings in rocky wooded bluffs, limestone glades,
ledges and tops of
limestone cliffs, and crevices of limestone cliffs. The bluffs and
cliffs are typically located along rivers (the Mississippi River in the
case of Illinois). This conservative species is found primarily in high
quality natural areas.
Faunal
Associations: Very little information is
available about floral-faunal relationships for Drummond's Goldenrod.
However, goldenrods (Solidago spp.) in general
attract a wide variety of insects to their flowerheads. This includes
honeybees, bumblebees, cuckoo bees (Epeolus spp.),
leaf-cutter bees (Megachile spp.), long-horned bees (Melissodes
spp.), little carpenter bees (Ceratina spp.),
green metallic bees (Augochlorella spp.), Halictid
bees (Halictus spp., Lasioglossum spp.),
plasterer bees (Colletes spp.), masked bees (Hylaeus
spp.), Andrenid bees (Andrena spp.),
dagger bees (Perdita spp., Heterosarus spp.),
ground beetles (Lebia spp.), ladybird beetles (Hippodamia
spp.), Sphecid wasps, Vespid wasps, Tiphiid wasps, Syrphid
flies, bee flies (Bombyliidae), Tachinid flies, flesh flies (Sarcophaga
spp.), blow flies (Lucilia spp.),
Muscid flies, butterflies, and day-flying moths (Robertson, 1929,
&
others). Other insects feed on the leaves, plant juices, and other
parts of goldenrods. The larvae of a leaf beetle, Microrhopala
excavata, have been observed to mine the leaves of Drummond's
Goldenrod. Other insect feeders of goldenrods include other leaf
beetles (Ophraella spp., Trirhabda spp.),
larvae
of leaf-mining flies (Calycomyza spp.), larvae of
gall flies (Asteromyia spp., Rhopalomyia
spp.), plant bugs (Lygus spp., Plagiognathus
spp., Slaterocoris spp.), stink bugs (Euschistus
spp.), aphids (Uroleucon spp.),
treehoppers (Stictocephala spp.), grasshoppers (Melanoplus
spp.), larvae of owlet moths (Cucullia spp.),
larvae of Geometer moths (various species), and larvae of Tortrix moths
(Epiblema spp., Eucosma spp., Phaneta
spp.).
Among vertebrate animals, the seeds of goldenrods are eaten by such
songbirds as the Eastern Goldfinch, Slate-colored Junco, Tree Sparrow,
and Indigo Bunting (Martin et al., 1951/1961; DeVore et al., 2004). The
White-tailed Deer and Cottontail Rabbit sometimes feed on the foliage
of these plants, while the Woodland Vole and other voles feed on both
the foliage and seeds (Martin et al., 1951/1961).
Photographic
Location: The wildflower garden of the webmaster
in Urbana, Illinois.
Comments:
This is one of the best goldenrods (Solidago spp.)
for ornamental
gardens. It should be cultivated more often. Drummond's Goldenrod
resembles many other goldenrod species, but it can be distinguished as
follows: 1) the leaves of Drummond's Goldenrod are unusually
wide, sometimes they are almost as wide as they are long, 2) the
leafy
bracts of the inflorescence of Drummond's Goldenrod are also unusually
wide, 3) its stems
are evenly short-pubescent, rather than hairless or hairy in
longitudinal lines, 4) the phyllaries, or floral bracts, of its
flowerheads have either bluntly acute or rounded tips, 5) the branches
of the terminal inflorescence spread rather widely, and 6) the habitats
of this goldenrod are largely restricted to riverside bluffs, limestone
glades, and
limestone cliffs. Of the preceding characteristics, #1 and #6 above are
the most distinctive. Some authors state that the leaves of this
goldenrod have 3 prominent veins (a central vein and 2 lateral veins) –
while this is true for some leaves, not all leaves have this
characteristic.