Description:
This wildflower is an annual about 2-6" tall. A typical plant has
a central stem that branches near the base, forming multiple
lateral stems about 2-12" long that are unbranched. These stems creep
along the ground or they are weakly ascending. Each stem is light green
to reddish brown, terete, and conspicuously hairy. Along the lower
one-third of each stem, pairs of opposite leaves are produced,
otherwise the leaves are alternate. Individual leaf blades are ¼-¾"
across and a little shorter in length; they are orbicular-reniform in
shape with 3-5 shallow rounded lobes or crenate teeth along their
margins. The leaf blades are medium green with appressed hairs above
and either appressed or spreading hairs below; their margins are
conspicuously ciliate, and they have a slightly succulent texture. The
petioles are the same length or a little shorter than the leaves, light
green, and
conspicuously hairy.
Individual flowers develop across from the
alternate leaves on slender pedicels about ¼-1" long. These pedicels
are light green and short-pubescent. Individual flowers are about 1/8"
across, consisting of a pale purple corolla with 4 spreading lobes, 4
green to reddish green sepals that are almost as long as the corolla, 2
stamens with
white anthers, and a 2-celled ovary with a single short style. The
corolla has several fine veins that originate from its center;
they are a darker shade of purple. The sepals are cordate-ovate in
shape, pubescent along their outer sides, and conspicuously ciliate
along their margins. The blooming period occurs from mid- to
late-spring for about 2 months; some plants may bloom
later in the
fall. The flowers are capable of self-pollination. Later, they are
replaced by 2-valved seed capsules about 1/8" (3 mm.)
across (or a little
more). Each seed capsule is globoid in shape, slightly flattened, and
only slightly constricted between the two valves; its exterior is
glabrous. Each capsule contains 4 seeds (2 seeds per cell). Each
small seed has a deep depression along one side. The root system
consists of a shallow branching taproot. This wildflower often forms
colonies of plants by reseeding itself.
Cultivation:
The
preference is full or partial sun, more or less mesic conditions, and
soil (possibly acidic) containing loam, sand, or gravel. This little
plant can spread aggressively by reseeding itself.
Range
&
Habitat: The non-native Ivy-Leaved Speedwell is uncommon
in southern
Illinois, while in the rest of the state it is rare or absent. It may
be more widespread than what is indicated by
these distribution records, however. This weedy wildflower was
introduced accidentally into North American from Europe. Habitats
consist of grassy slopes, fields, roadsides, and waste ground. Areas
with a history of disturbance are preferred.
Faunal
Associations: The nectar of the flowers attracts Andrenid
bees (
Andrena
spp.), Halictid bees (
Halictus
spp.,
Lasioglossum
spp.), and
miscellaneous Syrphid flies. Insects that feed on the foliage or suck
plant juices from
Veronica
spp. include the stink bug
Cosmopepla
lintneriana, the ebony bug
Corimelaena pulicaria,
and the flea beetle
Longitarsus turbatus.
The seeds are probably consumed by sparrows and
other birds, but there is a lack of records to substantiate this.
Cattle and other mammalian herbivores occasionally feed on the foliage
of these plants without apparent ill-effect.
Photographic
Location: On ground that was partially shaded
by a pine tree at
the Arboretum of the University of Illinois in Urbana, Illinois.
Comments: This
is one of several
Veronica
spp. from Europe that are weedy annuals
and they prefer similar habitats. Ivy-Leaved Speedwell can be
distinguished from similar species by a combination of the following
characteristics: 1) only non-terminal solitary flowers are
produced along its stems, rather than terminal or
non-terminal racemes, 2) the pedicels of its flowers (or seed capsules)
are about as long as the leaves, rather than shorter than the leaves,
3) the margins of its leaves have 3-5 blunt lobes or crenate
teeth, and 4) its stems, petioles, leaf margins, and sepal margins have
conspicuous long hairs, rather than short inconspicuous hairs or a lack
of hairs. This latter characteristic provides Ivy-Leaved Speedwell with
a bristly appearance. An
alternative spelling for the scientific name of this plant is
Veronica hederifolia.