Description:
This herbaceous plant is a summer annual about 1½–4' tall in Illinois,
although in tropical and subtropical areas it is a short-lived
perennial. The central stem is light green, 4-angled, and coarsely
hairy; it is unbranched below, but often becomes branched above. Pairs
of opposite leaves occur primarily along the lower central stem,
becoming more widely scattered above. The leaves are 1½-4" long and
¼-¾" across, becoming smaller as they ascend the stems; they are
oblong-oblanceolate or oblong-elliptic in shape and coarsely dentate
along their margins. The upper leaf surface is gray-green or
olive-green, wrinkled
along the veins, and nearly hairless, while the lower surface is pale
green and more hairy. The base of each leaf clasps the stem. The
central stem and ascending lateral stems terminate in panicled cymes of
flowers.
Each cyme consists of a dense flat-topped cluster of flowers
spanning about 1-3" across. Individual flowers are about 1/4" (5-6
mm.) across and 1/3" (8-9 mm.) long. Each flower consists of a tubular
pinkish purple corolla with 5 spreading lobes, a short tubular calyx
with 5 teeth, 4 inserted stamens, and a pistil with a single style. The
calyx is light green to reddish purple and hairy, while the lobes
of the corolla are slightly notched at their tips. Underneath the
flowers, are small lanceolate bractlets (about 1/8" or 3 mm. in
length). The
blooming period occurs from mid-summer into the fall for about 2-3
months. The flowers have a sweet fragrance. Each flower is replaced by
4 small nutlets that are narrowly oblongoid and flattened. This plant
reproduces by reseeding itself.
Cultivation:
The preference is
full sun and mesic to dry conditions. A variety of soil types are
readily tolerated. This plant can reseed itself aggressively in some
areas, although this appears to be less of a problem in Illinois than
in the southeastern states. Sometimes the leaves are discolored by
powdery mildew.
Range & Habitat: So far, the non-native
Purpletop Vervain has rarely escaped from cultivation and become
naturalized in Illinois (see
Distribution
Map). When wild plants do
occur, they rarely persist for very long. Purpletop Vervain was
introduced into the United States from Argentina and Brazil as an
ornamental garden plant. In Illinois, it has been encountered as a
naturalized plant in a weedy area along a pond among cattails (Coles
County), and it has been found growing wild at a construction site near
the Urbana Free Library by the webmaster (Champaign County). Outside of
the state, Purpletop Vervain has been found in such habitats as fields,
roadsides, weedy meadows, and other disturbed areas. It is often
cultivated in flower gardens.
Faunal
Associations: The flowers
of Purpletop Vervain are very attractive to nectar-seeking butterflies
and long-tongued bees. While little additional information about the
floral-faunal relationships of this wildflower is available, in general
Verbena spp.
(Vervains) are host plants of
Crambodes
talidiformis
(Verbena Moth), the aphid
Macrosiphum
verbenae, and the leaf beetle
Longitarsus suspectus
(Covell, 1984/2005; Thomas, 1877; Clark et al.,
2004). The seeds of these wildflowers are eaten in small amounts by
such songbirds as the Cardinal, Slate-Colored Junco, Field Sparrow,
Song Sparrow, Tree Sparrow, Swamp Sparrow, and White-Crowned Sparrow
(Martin et al., 1951/1961). Generally, the coarse and bitter
foliage is not attractive to mammalian herbivores.
Photographic Location: A construction site near the Urbana
Free Library in Urbana, Illinois.
Comments:
Purpletop Vervain is a lanky plant with brilliantly colored flowers
that are fragrant. Because these flowers attract the Monarch, Red
Admiral, Checkerspots, and other butterflies, it is a good choice for
the butterfly garden. While this species is regarded as invasive in
some areas of the United States, it appears to prefer highly disturbed
areas in Illinois, rather than high quality natural habitats. Sometimes
Purpletop Vervain is incorrectly referred to as 'Brazilian Vervain,'
which corresponds to another South American species,
Verbena
brasiliensis. Brazilian Vervain has escaped from
cultivation in
southeastern United States, but there are no records of this species
naturalizing in Illinois. Unlike Purpletop Vervain, Brazilian Vervain
produces distinct cylindrical spikes of flowers within each floral cyme
of the inflorescence (for a photographic comparison, see Brandenburg,
2010, p. 600).