Description:
This perennial wildflower is 2-5' tall and usually unbranched, except
where the inflorescence occurs. There are no basal leaves. The terete
central stem is light green, purplish red, or yellowish brown; the
typical variety is glabrous or sparsely short-pubescent, while var.
pubens
tends to be more pubescent. Alternate leaves occur along
the entire length of the stem; they are ascending and slightly
recurved. The lowermost leaves are small and scale-like, while the
remaining leaves are 3-5" long, ½-1" across, and relatively
uniform in size as they ascend the stem. The leaves are elliptic or
lanceolate-elliptic in shape, smooth and short-ciliate along their
margins, and either sessile or with short petioles. The upper leaf
surface is medium to dark green, while the lower surface is pale green
or whitish green. The upper and lower leaf surfaces of the typical
variety are glabrous to sparsely short-pubescent. The upper leaf
surface of var.
pubens
is sparsely to moderately short-pubescent or canescent, while
the lower surface is moderately to densely short-pubescent or
canescent.
The central stem terminates in a flat-headed panicle
(compound corymb) of flowerheads about 3-12" across. Individual
flowerheads are about ½" across, consisting of 5-12 ray florets that
surround 12-25 disk florets. The petal-like corollas of the ray florets
are white; the tubular corollas of the disk florets are yellow while in
bloom, but become cream-colored or dingy white thereafter. Each disk
floret has 5 spreading lobes. At the base of each flowerhead, there are
2-4 series of appressed floral bracts (phyllaries) that are narrowly
oblong, green, and glabrous to short-pubescent. The branches of the
inflorescence are light green or yellowish brown; they are either
glabrous or short-pubescent. Leafy bracts about ½-1½" long occur along
these branches; they are lanceolate to narrowly elliptic. The blooming
period occurs from late summer into the fall, lasting 1-2 months. Both
ray and disk florets are replaced by achenes with whitish tufts of
hair. In each tuft of hair, the outermost hairs are bristly and short
(less than 1 mm. in length), while the inner hairs are longer (3-6 mm.
in length). Individual achenes are about 3 mm. (1/8") long,
bullet-shaped, and sparsely short-pubescent. They are distributed by
the wind. The root system is fibrous and rhizomatous. Occasionally,
small colonies are formed from vegetative offsets.
Cultivation:
The preference is full or partial sun, slightly wet to moist
conditions, and calcareous soil that contains sandy-loam. The pH
should be slightly acidic. This wildflower can adapt to other kinds of
soil, although they are not preferred.
Range
& Habitat: The
native Flat-Topped Aster is occasional in NE Illinois and parts of
central Illinois, while in the rest of the state it is uncommon or
absent. Habitats include wet to moist sand
prairies, moist sandy thickets, soggy meadows and openings in wooded
areas, interdunal sloughs and swales near Lake Michigan, fens, and
seeps. This wildflower is found in higher quality wetlands that are
often sandy and calcareous.
Faunal
Associations:
The nectar and pollen of the flowerheads
attract
honeybees, bumblebees, little carpenter bees (Ceratina spp.),
long-horned bees (Melissodes spp.), leaf-cutting bees (Megachile spp.),
cuckoo leaf-cutting bees (Coelioxys spp.), Halictid bees, Andrenine
bees, dagger bees (Perdita spp., Pseudopanurgus spp.), Sphecid wasps,
Tiphiid wasps, Eumenine wasps, paper wasps (Polistes spp.), hornets
(Dolichovespula spp., Vespula spp.), ants, Syrphid flies, bee flies
(Bombyliidae), skippers, and beetles (Grundel et al., 2011; Wilhelm
& Rericha, 2017). Oligolectic bees (specialist pollinators) that
have been observed to visit Flat-topped Aster (Doellingeria umbellata),
are
Andrena nubecula,
Andrena placata,
Perdita octomaculata, and
Pseudopanurgus andrenoides.
The foliage, plant sap, roots, and other
parts of Flat-topped Aster and other asters are eaten by such insects
as leaf beetles, weevils, larvae of tumbling flower beetles, larvae of
leaf-miner flies, larvae of gall flies, larvae of fruit flies, plant
bugs, aphids, leafhoppers, armored scales, mealybugs, larvae of moths,
larvae of crescent and checkerspot butterflies (Phyciodes spp.,
Chlosyne spp.), and grasshoppers. Flat-topped Aster is the primary host
plant of the butterfly, Harris' Checkerspot
(Chlosyne harrisii). The
Insect Table
has a more detailed list of these insect feeders. The seeds of asters
are eaten by the American Tree Sparrow, while the Ruffed Grouse feeds
on both the leaves and seeds (Martin et al., 1951/1961). Hoofed
mammalian herbivores frequently browse on these plants, particularly
the less hairy species. These species include Elk, White-tailed Deer,
cattle, and other domesticated grazers (Schneider et a., 2006; personal
observations).
Photographic
Location: A wet sand prairie at the Indiana Dunes National
Lakeshore in NW Indiana.
Comments: Flat-Topped Aster is an attractive wildflower
that favors
relatively open areas that are damp and sandy. Because of its
flat-topped
inflorescence, it is relatively easy to distinguish from other asters
(
Aster spp.)
within the state. In particular, the prominent
disk florets of the flowerheads are somewhat unusual in that they
become dingy white or cream-colored shortly after they bloom, rather
than orange-red or purple. The reason Flat-Topped Aster has been
assigned to the
Doellingeria
genus is related to the different hair
lengths of its tufted achenes: the outer hairs are less than 1 mm. in
length, while the inner hairs are 3-6 mm. in length. Usually, asters
have a uniform length for the hairs of their tufted achenes. An older
scientific
name of Flat-Topped Aster is
Aster
umbellatus.