Description:
This herbaceous perennial plant is ¾–2¼' tall and unbranched. The
central stem is green, glabrous, and terete. A single well-developed
leaf occurs near the base of the central stem. This leaf is 4-12" long,
¼–1½" across, and ascending; it is linear-elliptic to elliptic in shape
and entire (toothless) along its margins. The leaf tapers gradually
into a narrow base that is enclosed by a sheath, while its tip is
narrowly acute and hull-shaped. The upper surface of the
entire leaf is often slightly concave (curved inward from the margins)
along its length. Both the upper and lower leaf surfaces are yellowish
green to medium green and glabrous. Leaf venation is parallel. Below
the well-developed leaf, there are 1-2 rudimentary leaves that are
sheath-like and inconspicuous.
The central stem terminates in a spike
of 2-15 flowers (rarely up to 20). The rachis of this floral spike is
green to reddish purple, terete, and glabrous; it has a tendency to
zig-zag between the flowers. Each flower is 1-2" across, consisting of
3 petaloid sepals, 3 petals, an exposed reproductive column, and an
inferior ovary. Both the sepals and petals (excluding the petal
that has been modified into a lip) are pink to deep rosy pink (rarely
white); the sepals and petals may also have faint veins of dark rosy
pink. The sepals are ovate or broadly oblong-ovate in shape, while
the petals are broadly elliptic or elliptic-ovate in shape. Depending
on the stage of development, both petals and sepals are spreading and
more or less incurved from their tips; their upper surfaces are flat to
somewhat concave. Both sepals and petals are nearly the same length
(about ¾" long), although the sepals are a little wider. The third
petal has been modified into an upper lip about ¾" long. This upper lip
is linear in shape (grooved above and convex below), except toward its
tip, where it has been widened by 2 lateral lobes; these lobes are
half-orbicular to bluntly triangular in shape. At the center of this
lobed tip, is a patch of white and a cluster of clubbed pseudo-stamens;
these pseudo-stamens are hair-like in appearance and yellow to orange.
Elsewhere, the lip is pink to rosy pink (rarely white) like the sepals
and
remaining petals. The lip is also hinged at its base.
The exposed
reproductive column is mostly linear-flattened in shape, straight, and
pink to rosy pink (rarely white). However, toward its tip, the
reproductive column is upturned, terete along its center, and laterally
lobed. The reproductive organs are located at the tip of the
column, which is dark rosy pink. The reproductive column lies opposite
from the upper lip of the flower; it is about ¾" long. The ascending
inferior ovary, at the time of bloom, is light green to pale greenish
pink, cylindrical-ribbed in shape, and glabrous. The blooming
period usually occurs from early to late summer, lasting about 3-4
weeks. The flowers bloom sequentially from the bottom to the top of the
floral spike. There can be either a mild floral fragrance or none.
Afterwards, fertile flowers are replaced by seed capsules that are
about ¾" long and ellipsoid-ovoid in shape; they eventually break open
to release numerous tiny seeds, which are distributed by the wind. The
root system consists of a globoid to ovoid corm with fibrous roots
below.
Cultivation:
The preference is full or partial sun, wet
to moist conditions, and soil containing some sand and/or peat.
Sometimes, this orchid is available from specialist orchid nurseries
and it can be cultivated in gardens if its requirements are met. Wild
plants, however, should never be collected.
Range & Habitat:
The native Grass Pink Orchid has been found primarily in the northern
half of Illinois and a few scattered counties elsewhere (see
Distribution
Map).
This orchid is now rare and it is state-listed as
'endangered.' At one time, the Grass Pink Orchid was more abundant in
Illinois, but its populations have declined because of habitat
destruction and unscrupulous collecting. Habitats where this orchid can
be found include wet to moist sand prairies, wet to moist sandy
meadows, shallow sandy swales, openings in sandy swamps, fens, and
bogs. This orchid is found in high quality natural areas.
Faunal
Associations: Bumblebees and other large
long-tongued bees are the
primary pollinators of the flowers. Halictid bees,
flies, butterflies, skippers, and beetles also visit the flowers
occasionally, but they are unlikely to be effective at
cross-pollination. See Cole (1988), Luer (1975), Small (1976), and
Thien & Marcks (1972) for more information. Neither nectar nor
accessible pollen are available to such flower-visiting insects.
Instead, they are lured by deception to land on the showy flowers. In
particular, they are often attracted to the colorful pseudo-stamens on
the lip of the flower. If the visiting insect has sufficient weight,
the hinged lip of the flower collapses onto its exposed reproductive
column, attaching pollinia to the back of the insect. When the same
insect visits the next flower, the same process can deposit the
pollinia onto the exposed stigma of the reproductive column, enabling
cross-pollination to occur. Like other orchids, the foliage and flowers
of the Grass Pink Orchid are probably vulnerable to browsing by
White-tailed Deer and other mammalian herbivores. When such animals are
too abundant, it may be necessary to protect colonies and solitary
plants of this orchid with wire cages or fencing.
Photographic Location:
A shallow sandy swale at a nature preserve in
Lake County, Illinois.
Comments:
The structure of the flower for this orchid is highly unusual because
its lip is located at the top rather than the bottom, causing the
flower to appear upside-down (even though it is actually rightside-up).
The Grass Pink Orchid is the most common species of its genus within
the state and it is widely distributed within the eastern United States
and SE Canada. The only other species of this genus within the state,
the Oklahoma Grass Pink Orchid (
Calopogon oklahomensis),
is even more
rare and it is also state-listed as 'endangered.' This latter orchid
can be distinguished from the former as follows: 1) the petals and
sepals of its flower are usually a lighter shade of pink, and 2) the
lip of its flower has a patch of short pink pseudo-stamens above its
showier yellow pseudo-stamens. The more common Grass Pink orchid lacks
short pink pseudo-stamens on the lip of its flower. The Oklahoma Grass
Pink Orchid is also found in drier habitats, like mesic prairies. Other
common names of
Calopogon tuberosus are Tuberous
Grass Pink and Grass
Pink, even though it is neither a grass nor a member of the Pink family
(Caryophyllaceae).