Description:
This perennial plant is 1-2½' tall and unbranched. The central stem is
light green, 4-angled, and glabrous to short-pubescent. Pairs of
opposite leaves are distributed rather abundantly along the central
stem. Individual leaves are 1½-4" long and ½-1½" across; they are
elliptic to broadly elliptic in shape and coarsely dentate along their
margins. There is typically 5-7 large teeth along each side of a leaf.
Each leaf tapers to a wedge-shaped base and it tapers gradually to an
acute tip. The upper leaf surface is medium to dark green and hairless
(or nearly so), while the lower surface is pale
green and hairless to
short-pubescent. Sometimes, the leaves become purplish green in bright
sunlight. Dense whorled clusters of nearly sessile flowers occur above
the axils of the middle to upper leaves. Each flower is about 1/8" (3
mm.) in length; it consists of light green calyx with 4-5 teeth, a
white corolla with 4-5 tiny lobes, 2 slightly exerted stamens, and a
pistil. The toothed calyx is 0.5-1 mm. in length; its teeth are broadly
triangular. The corolla is 2.5-3.5 mm. in length; it is campanulate
(bell-shaped). The blooming period occurs from mid-summer into the
fall, lasting 2-3 months. Usually, only a few flowers are in bloom at
the same time. Afterwards, each flower is replaced by 4 nutlets. The
nutlets are 1.0-1.5 mm. long, obovoid, and 3-angled; the apex of each
nutlet is tuberculate (bumpy) along the outer side, while
the inner sides are flat and depressed. Taken together, the upper
surface of each group
of 4 nutlets forms an indented central area and a bumpy
exterior. The root system
is stoloniferous and tuberous. Clonal colonies of plants are often
formed.
Cultivation:
The preference is full or partial sun, wet to moist
conditions, and soil that is sandy, loamy, or marly.
Range
& Habitat: The native Northern Bugleweed is
occasional in the
northern half of Illinois, while in the southern half of the state it
is rare or absent (see
Distribution
Map). Generally, Northern Bugleweed
has a more northern distribution in North America than most species in
its genus. Habitats include marshes, fens, low areas along pools of
water, streambanks, and swamps. These habitats include both sandy and
non-sandy wetlands. Should a wildfire occur, this wildflower can
regenerate itself through the stolons and tubers of its root system.
Faunal
Associations: The small flowers are cross-pollinated by a
variety of
insects, including long-tongued bees, short-tongued bees, wasps, and
flies. Most of these floral visitors suck nectar, although some bees
and flies may use the pollen as a food source. Small butterflies and
skippers are less typical visitors of the flowers. Among vertebrate
animals, muskrats consume the tubers of Northern Bugleweed and other
Lycopus spp.
with tuberous root systems. Generally, the foliage is not
attractive to mammalian herbivores because of its bitter taste. In
addition, the foliage is probably somewhat toxic to them if it is eaten
in quantity.
Photographic Location: Along a sandy marsh at the Heron
Boardwalk in
Vermilion County, Illinois.
Comments:
Northern Bugleweed is a fairly typical example of the
Lycopus spp. that
occur in Illinois. Its calyx can be distinguished from those of similar
species by its short length (1.0 mm. or less) and broad
teeth. Other
Lycopus
spp. within the state have longer calyces
with narrow teeth. An exception is
Lycopus
virginicus
(Virginia
Bugleweed), which has the same type of calyx. This latter species,
however, has leaves that taper abruptly toward their bases, rather
than gradually, and its leaf margins have more abundant smaller teeth.
The root systems of these two species are also somewhat different:
Northern Bugleweed usually produces tubers, while Virginia Bugleweed
lacks such tubers.