Description:
This
perennial wildflower consists of a small rosette of basal
leaves about 3-8" across, persisting from autumn to spring. These basal
leaves are semi-evergreen, ¾-2½" long, and ½-2" across; they are
oval-cordate in shape and their margins are crenate-serrate. The upper
surfaces of basal leaves are medium green and sparsely to
moderately short-pubescent, while their lower surfaces
are light
green and moderately to densely short-pubescent. Exposure to
near-freezing temperatures sometimes causes the basal leaves to turn
purple. The petioles of basal leaves are up to 2" long,
slender,
and short-pubescent. This wildflower bolts during the late spring to
produce one or more leafy stems about 1½-3¼' long that are initially
erect or ascending, although tending to sprawl across the ground later
in the year. These stems are light green to purple, terete, and
short-pubescent to pubescent throughout; this pubescence is not
restricted to longitudinal lines. The stems are unbranched below, while
they are sparingly to occasionally branched above.
Alternate leaves
occur along these stems that are 1-4" long and ½-3" across (becoming
smaller in size as they ascend the stems); they are cordate below,
becoming cordate-ovate to lanceolate above. The margins of these
alternate leaves are coarsely serrated below, becoming serrated to
slightly crenate-serrate above. The upper surfaces of these leaves are
moderate green and sparsely short-pubescent to densely short-pubescent,
while their lower surfaces are light green and moderately to densely
short-pubescent or pubescent throughout; this pubescence is not
restricted to the major veins. When these leaves are exposed to bright
sunlight, they may become yellowish green. Leaf texture tends to be
stiff and rough. The petioles of alternate leaves are ½-2" long and
their margins
are more or less winged; they are moderately to densely
short-pubescent. Sometimes small secondary leaves appear in the axils
of these leaves. Each leafy stem terminates in a
panicle of flowerheads about ½-2' long and about one-half as much
across. In addition to the central stalk of the panicle, there
are several ascending lateral branches along
which most of the flowerheads occur on short peduncles (less than ½"
long). The central stalk, branches and peduncles are light green to
purple, terete, and evenly short-pubescent to pubescent.
Each
flowerhead is about ½" across, consisting of 10-15 ray florets that
surround 10-15 congested disk florets. The petaloid rays of these
flowerheads are
usually lavender or light violet (less often white) and linear-oblong
in shape. The short-tubular disk florets are initially cream-colored to
yellow, but they later become reddish purple. Each disk floret has 5
short lobes that are erect. Around the base of each flowerhead, the
scale-like phyllaries (floral bracts) are arranged in several
overlapping series; these phyllaries are appressed together, or nearly
so. Individual phyllaries are 4-6 mm. in length, linear-lanceolate in
shape, and light green, except for diamond-shaped patches of dark green
near their apices. Leafy bracts occur along central stalk and lateral
branches of the panicle
that are up to 1" long and elliptic to linear-oblong in shape; they are
slightly toothed to entire (smooth) along their margins. The blooming
period occurs from late summer into autumn for 1-3 months.
Afterwards, the florets are replaced by achenes with small tufts of
white hair. The achenes are 2-3 mm. long, oblongoid-oblanceoloid in
shape, slightly ribbed, and light to medium brown (sometimes with
reddish or
purplish tints). They are distributed by the wind. The root system
is fibrous and short-rhizomatous. On older plants, a small woody caudex
may develop. Clonal offsets are often produced from the rhizomes.
Cultivation:
The
preference is partial sun, mesic to dry-mesic conditions, and soil
containing loam, clay-loam, or some rocky material. This wildflower is
usually easy to cultivate. The foliage may become discolored with age;
lower leaves may wither away in response to hot dry conditions. There
is a tendency for the flowering stems to sprawl across the ground
during the autumn.
Range
& Habitat: The native Drummond's
Aster is common in central and northern Illinois, while in the southern
section of the state it is occasional to locally common (see
Distribution
Map).
Habitats
include upland woodlands,
rocky woodlands, woodland openings, woodland borders, areas along
woodland paths, disturbed open woodlands, upland savannas, thinly
wooded bluffs, powerline clearances through wooded areas, thickets,
partially shaded riverbanks, burnt-over vacant lots, edges of yards,
and partially shaded areas along roadsides. Drummond's Aster can be
found in both high quality natural areas and degraded sites. Occasional
wildfires are beneficial when they reduce excessive shade from overhead
canopy trees. The woodlands where this aster occurs are dominated by
various deciduous trees, especially oaks (
Quercus spp.).
Faunal Associations: The flowers are
cross-pollinated by honeybees,
bumblebees, Halictid bees (including green metallic
bees), Andrenid bees, miscellaneous wasps, Syrphid flies,
Tachinid flies, Muscid flies, butterflies, skippers, moths,
and other insects. Both nectar and pollen are available as floral
awards to
such visitors. The following bees are oligolectic visitors (pollinator
specialists) of aster flowers:
Andrena
asteris,
Andrena
asteroides,
Andrena
hirticincta,
Andrena
nubecula,
Andrena
solidaginis, and
Colletes
simulans armata. Some of these oligoleges also visit
goldenrod flowers. Other insects that are associated with Drummond's
Aster and other
asters (
Symphyotrichum
spp.)
include leaf beetles, larvae of fruit flies and
leaf-mining flies, plant bugs, lace bugs, stink bugs, aphids,
leafhoppers, walkingsticks, caterpillars of butterflies, and
caterpillars of many moths. These insects feed destructively on the
leaves, flowers, developing seeds, stems, roots, or plant
juices (see the
Insect Table and
Moth
Table for more information about
these species). In general, asters are less important to vertebrate
animals. However, such upland gamebirds as the Ruffed Grouse and Wild
Turkey feed occasionally on the foliage and seeds; such mammals as
deer, rabbits, groundhogs, and livestock also feed on the foliage.
Photographic Location: Edge of a wooded
area at Crystal Lake Park in
Urbana, Illinois.
Comments:
Drummond's Aster (
Symphyotrichum
drummondii) is part of a complex of similar
species involving Arrow-Leaved Aster (
Symphyotrichum sagittifolium),
Heart-leaved Aster (
Symphyotrichum
cordifolium), and White Arrowleaf Aster
(
Symphyotrichum
urophyllum).
There has been a history of taxonomic instability
involving the interrelationships between these 4 species. Some of these
species are considered varieties of each other by some taxonomists, or
they have been merged with other species within this complex.
Drummond's Aster can be distinguished from other species (or varieties)
in this complex by the evenly distributed pubescence along its stems
and by the evenly distributed pubescence on the lower sides of its
leaves. The abundance and conspicuousness of this pubescence can vary
across different populations of plants in Illinois. Drummond's Aster
also differs from Heart-leaved Aster by its winged petioles; this is a
trait that it shares with the remaining species of this complex.
Relatively recently, all of these species have been assigned
to the
Symphyotrichum
genus; Drummond's Aster was referred to previously as
Aster
drummondii. While the individual
flowerheads are relatively small, the inflorescence of this aster is
often quite large and showy during the autumn.