Description:
This perennial wildflower is ¾-3' tall and usually unbranched, except
near the apex where the inflorescence occurs. Erect leafy stems may be
produced individually or in small clusters. The central stem is is
whitish green, 4-angled, and covered with either glandular or
non-glandular hairs (or both). Pairs of opposite leaves occur along
each stem. The leaf blades are 1-4" long and ¾-3" across; they are
cordate in shape with coarse dentate-crenate margins. The upper blade
surface is medium to dark green and either sparsely covered with short
hairs or glabrous. The lower surface is slightly more pale and varies
from densely hairy to sparsely hairy (rarely glabrous). The slender
petioles are ½-2" long and hairy.
The central stem terminates in a raceme of flowers about 3-12" long. In
addition, any upper lateral
stems may produce smaller racemes of flowers. The racemes are narrow
and usually only a few flowers are in bloom at the same time.
Individual flowers are ½-1" long. Each flower consists of a
blue-violet or purple corolla with 2 lips, a
short tubular calyx with 2 lips, 4 inserted stamens, and
a pistil with an inserted style. The corolla is much longer than the
calyx. The corolla is narrow and tubular at its base, but becomes more
wide and swollen towards its lips. The upper lip of the corolla
consists of a protective hood, while the lower lip is larger, more
rounded, and somewhat lobed. There is usually a large patch of white
along the inside of the
lower lip that is speckled blue-violet or purple. The calyx is about
1/8" (3 mm.) long, whitish
green, and covered with glandular hairs. A protuberance occurs along
the upper side of each calyx. The pedicels of the
flowers are very short
(about 1/8" or 3 mm. in length). Underneath the flowers, there are
leafy
bracts of variable length: on some plants, they extend as far as
the tips of the calyces, but no further, while on other plants they
extend beyond the tips of the calyces. These bracts are
ovate in shape. The blooming period typically occurs from late spring
to mid-summer, lasting about 1 month. However, if a stem is
damaged, this may delay
flowering until later in the year. Afterward, the flowers are replaced
by small dark nutlets. The root system is fibrous and either
rhizomatous or
stoloniferous.
Cultivation:
The preference is dappled sunlight to medium shade, mesic
conditions, and soil that contains loam, some calcareous sand,
or rocky material (typically limestone). This wildflower is a good
choice for shade gardens.
Range
& Habitat: The native Heart-Leaved Skullcap is
occasional throughout Illinois. Habitats include
upland woodlands that are often rocky, bottomland woodlands,
bluffs, woodland openings, shaded areas along cliffs, edges of
limestone glades, and thickets. This wildflower is usually found in
higher quality woodlands where the original ground flora is still
intact.
Faunal
Associations: The flowers are cross-pollinated by
long-tongued
bees, particularly bumblebees and Anthophorine bees (
Anthophora spp.),
which suck nectar and collect pollen. Short-tongued bees and Syrphid
flies may also visit the flowers, but they are too small to be
effective pollinators. Swallowtail butterflies and other butterflies
occasionally visit the flowers, but they are also less effective at
cross-pollination according to Robertson (1929). A few species of
insects feed on the leaves of Heart-Leaved Skullcap and other
skullcaps (
Scutellaria
spp.). These species include the skeletonizing
leaf beetle
Phyllobrotica
limbata,
Asphaera
lustrans (Shiny Flea
Beetle), larvae of
Prochoreutis
inflatella (Skullcap Skeletonizer
Moth), and larvae of the moth
Caloptilia
scutellariella. The larvae of
this last species are both blotch leaf-miners and leaf-folders. Because
of its bitter taste and possible toxicity, mammalian
herbivores rarely feed on the foliage.
Photographic
Location: A floodplain woodland in Lake County, Illinois.
The
photographed plants are either
Scutellaria
ovata ovata or
Scutellaria
ovata bracteata. The photographs (Copyright © 2011) were
taken by Paul Showers.
Comments: Unlike many other
Scutellaria spp.,
this skullcap has mostly heart-shaped leaves with indented bases;
sometimes the uppermost leaves have bases that are rounded, rather than
indented. Across its range, Heart-Leaved Skullcap is highly
variable and several subspecies have been recognized. In Illinois,
three of these subspecies can be found:
Scutellaria ovata ovata,
Scutellaria ovata
bracteata, and
Scutellaria
ovata rugosa.
The first two subspecies occur throughout Illinois and they are
distinguished by the size of the leafy bracts on their racemes:
Scutellaria ovata ovata
has leafy bracts that extend no farther than the calyces of the
flowers, while
Scutellaria
ovata bracteata has leafy bracts that extend beyond the
calyces of the flowers. The third subspecies,
Scutellaria ovata rugosa,
is only found in southern Illinois. It can be distinguished from the
preceding subspecies by its small size (less than 1' tall) and small
leaves (less than 1½" long).